NATO spokeswoman Oana Lungescu said Friday that "Russian statements threatening to target allies are unacceptable and counterproductive."
Putin said Thursday that Moscow has tested an array of new strategic nuclear weapons that can't be intercepted, telling the West: "You have failed to contain Russia."
Lungescu said NATO's missile defense system is built to respond to attacks from outside Europe and North America and not directed against Russia.
Noting Russia's "aggressive actions" in Ukraine and military buildup around Europe, she said: "NATO is pursuing a twin-track approach to Russia: strong deterrence and defense, combined with meaningful dialogue."
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